不同播栽方式下杂交籼稻茎秆生长和穗粒形成特点及与气象因子的关系

doi:10.16819/j.1001-7216.2016.6012

摘要/Abstract

摘要：

为探明不同播栽方式下杂交籼稻幼穗分化期茎秆和幼穗生长的规律及差异以进一步明确不同播栽方式穗粒形成特点,于2014年采用两因素裂区试验设计,研究了机直播、机插和手插3种播栽方式下2个不同穗粒型杂交籼稻组合[宜香优2115(中穗型)和F优498(大穗型)]的穗分化期茎秆和幼穗生长规律和穗粒形成特点,并分析了幼穗分化期气象因素与幼穗和茎秆生长的关系。结果表明:1)基部向上第1至第3伸长节间长度分别在抽穗前20 d、16 d、12 d后趋于稳定,故基部第1、2节间的降长增粗应在抽穗前16 d之前;穗干质量和穗茎比在抽穗前12 d后迅速增加。2)从抽穗前16 d开始至抽穗期,穗茎干重比与多数枝梗和颖花性状呈显著或极显著正相关,穗与茎秆竞争同化物的能力直接影响到穗粒形成。3)机插穗分化中后期穗茎干质量比较高,且抽穗期穗干质量显著高于手插和机直播,有利于提高其每穗粒数,且机插拔节后群体生长率及单茎和群体干物质积累量较高,抽穗期叶面积指数较高,粒叶比高于机直播,较机直播更利于粒重的提高和穗长及着粒数的增加;大穗型品种F优498抽穗期穗干质量显著高于中穗型品种宜香优2115,且穗分化后期穗茎干质量比高于宜香优2115,是其每穗粒数高的重要原因;F优498穗部着粒较密,较大的叶面积指数及粒叶比利于大穗的形成。4)机插全生育期最长,机直播最短,且机插穗分化期历时较长,穗分化期积温和日照时数较高,为穗粒形成提供了较好的温光条件;茎秆长度和干质量及穗长受气象因子影响较大,孕穗前适宜的温度和一定的积温有利于茎秆和幼穗伸长及茎秆的物质充实。生产上应针对不同播栽方式及品种的生育进程差异,适时采取措施调节茎秆和幼穗生长及物质分配,在培育壮秆的基础上提高穗分化期穗茎比以达到增粒增产的目的。

关键词: 水稻, 穗, 茎秆, 穗粒形成, 机插, 机直播, 气象因子

Abstract:

The objective of this study is to explore the growth pattern and differences of stem and panicle of indica hybrid rice under various planting methods in panicle differentiation stage, and to investigate the effects of planting methods on grain formation. A split plot experiment was carried out in 2014 with two factors, including variety (Yixiangyou 2115 and F you 498) and planting method (mechanized direct-seeding, MD; mechanized transplanting, MT; hand transplanting, HT).The Growth pattern of stem and panicle, the formation characteristics of grain were researched, and the relationship between meteorological factors in panicle differentiation stage and the growth of stem and panicle were analyzed. The results were as follows: 1) Length of first to third elongated internodes from the base stablized 20 d, 16 d and 12 d before heading,respectively, indicating that reducing length and increasing roughness of basal first and second internodes should be done 16 d before heading. Dry panicle weight and ratio of panicle to stem increased rapidly 12 d before heading. 2) Dry weight ratio of panicle to stem was significantly positively correlated with rachis branches and spiketets from 16 d before heading to heading date. The ability that panicle competing assimilates with stem directly affected the formation of grain. 3) MT had higher dry weight ratio of panicle to stem in middle and late differentiation stage of panicle, MT also had higher dry matter accumulation of panicle than MD and HT in heading stage. So MT had an advantage in the number of grains. MT also had higher crop growth rate of population and higher dry matter accumulation per stem and population than MD and HT after jointing. MT also had higher leaf area index in heading, and its grain leaf ratio was higher than MD, so MT contributed to more grains and spikelets, longer panicle length. F you 498 with large panicle had higher dry matter weight of panicle in heading than Yixiangyou 2115 with middle panicle, and it had higher dry matter weight ratio of panicle to stem than Yixiangyou 2115, these were the important reason for that F you 498 had higher number of grains. Denser grain of F you 498 compared to Yixiangyou 2115 resulted from its larger leaf area index and grain to leaf ratio.4) MT had the longest growth duration,while MD had the shortest. MT had a longer panicle differentiation stage, higher accumulated temperature and illumination hours, providing a good temperature and light condition for the formation of grain in MT. Length and dry weight of stem and length of panicle were greatly influenced by meteorological factors. Suitable temperature and a certain amount of accumulated temperature before booting was conducive to the elongation of stem and panicle and filling of stem. Given the differences in growth under different planting methods and of varieties, we should take measures to regulate growth of stem and panicle and the distribution of assimilates in an appropriate time. In order to achieve the goal of increasing grain yield, the dry weight ratio of panicle to stem should be increased on the basis of cultivating strong stalk.

Key words: rice, panicle, stem, formation of grain, mechanized transplanting, mechanized direct-seeding, meteorological factors.

中图分类号:

S511.01

田青兰, 刘波, 孙红, 何莎, 钟晓媛, 赵敏, 任万军. 不同播栽方式下杂交籼稻茎秆生长和穗粒形成特点及与气象因子的关系[J]. 中国水稻科学, 2016, 30(5): 507-524.

Qing-lan TIAN, Bo LIU, Hong SUN, Sha HE, Xiao-yuan ZHONG, Min ZHAO, Wan-jun REN. Characteristics of Stem Growth and Formation of Grain of indica Hybrid Rice in Different Planting Methods and Their Correlation with Meteorological Factors[J]. Chinese Journal OF Rice Science, 2016, 30(5): 507-524.

图/表 16

参考文献 35

[1]	丁颖, 李乃铭, 徐雪宾. 水稻幼穗发育和谷粒充实过程的观察. 农业学报, 1959, (10): 59-85.
	Ding Y, Li N M, Xu X B.Observation of panicle development and courses of gain of rice.Agric Sci, 1959, (10): 59-85. (in Chinese with English abstract)
[2]	刘利, 雷小龙, 王丽, 等.种植方式对杂交稻枝梗和颖花分化及退化的影响. 作物学报, 2013, 39(8): 1434-1444.
	Liu L, Lei X L, Wang L, et al.Effect of planting method on differentiation and retrogression of branches and spikelets of hybrid rice cultivar.Acta Agron Sin, 2013, 39(8): 1434-1444. (in Chinese with English abstract)
[3]	娄伟平, 孙永飞, 张寒, 等. 温度对每穗颖花数的影响. 浙江农业学报, 2005, 17(2): 101-105.
	Lou W P, Sun Y F, Zhang H, et al.Effects of temperatures on spikelets per panicle of rice.Acta Agric Zhejiangensis, 2005, 17(2): 101-105. (in Chinese with English abstract)
[4]	曾研华, 张玉屏, 向镜, 等. 籼型常规早稻穗分化期低温对颖花形成和籽粒充实的影响. 应用生态学报, 2015, 26(7): 2007-2014.
	Zeng Y H, Zhang Y P, Xiang J, et al.Effects of low temperature on formation of spikelets and grain filling of indica inbred rice during panicle initiation in early-season.Chin J Appl Ecol, 2015, 26(7): 2007-2014. (in Chinese with English abstract)
[5]	李刚华, 王惠芝, 王绍华, 等. 穗肥对水稻穗分化期碳氮代谢及颖花数的影响. 南京农业大学学报, 2010, 33(1): 1-5.
	Li G H, Wang H Z, Wang S H, et al.Effect of nitrogen applied at rice panicle initiation stage on carbon and nitrogen metabolism and spikelets per panicle.J Nanjing Agric Univ, 2010, 33(1): 1-5. (in Chinese with English abstract)
[6]	王惠芝, 尤娟, 王绍华, 等. 迟熟中粳稻穗茎生长与每穗颖花数的关系. 作物学报, 2007, 33(5): 820-825.
	Wang H Z, You J, Wang S H, et al.Relationship between panicle and stem growth and spikelets per panicle in late-maturing medium japoncia rice.Acta Agron Sin, 2007, 33(5): 820-825. (in Chinese with English abstract)
[7]	杨世民, 谢力, 郑顺林, 等. 氮肥水平和栽插密度对杂交稻茎秆理化特性与抗倒伏性的影响. 作物学报,2009, 35(1): 93-103.
	Yang S M, Xie L, Zheng S L, et al.Effects of nitrogen rate and transplanting density on physical and chemical characteristics and lodging resistance of culms in hybrid rice.Acta Agron Sin, 2009, 35(1): 93-103. (in Chinese with English abstract)
[8]	李国辉, 钟旭华, 田卡, 等. 施氮对水稻茎秆抗倒伏能力的影响及其形态和力学机理. 中国农业科学, 2013, 46(7): 1323-1334.
	Li G H, Zhong X H, Tian K, et al.Effect of nitrogen application on stem lodging resistance of rice and its morphological and mechanical mechanisms.Sci Agric Sin, 2013, 46(7): 1323-1334. (in Chinese with English abstract)
[9]	陈小荣, 钟蕾, 贺晓鹏, 等. 稻穗枝梗和颖花形成的基因型及播期效应分析. 中国水稻科学, 2006, 20(4): 424-428.
	Chen X R, Zhong L, He X P, et al.Effects of genotype and seeding-date on formation of branches and spikelets in rice panicle.Chin J Rice Sci, 2006, 20(4): 424-428. (in Chinese with English abstract)
[10]	曾研华, 张玉屏, 王亚梁, 等. 籼粳杂交稻枝梗和颖花形成的播期效应. 中国农业科学, 2015, 48(7): 1300-1310.
	Zeng Y H, Zhang Y P, Wang Y L, et al.Effect of sowing date on formation of branches and spikelets in indica japonica hybrid rice.Sci Agric Sin, 2015, 48(7): 1300-1310. (in Chinese with English abstract)
[11]	凌启鸿. 水稻茎秆维管束数与穗部性状关系及其应用的研究. 江苏农学院学报, 1982, (3): 48-58.
	Ling Q H.The relationship between the number of macro-vascular bundles in culm and the panicle characters in rice plant and its application.J Jiangsu Agric Coll, 1982,(3): 48-58. (in Chinese with English abstract)
[12]	苏祖芳. 水稻单茎茎鞘重与产量形成关系及其高产栽培途径的探讨. 江苏农学院学报, 1993, 14(1): 37-39.
	Sun Z F.Discussion on the relationship between the culm-sheath weight per shoot and yield in rice and its cultural approaches of high yield.Acad J Jiangsu Agric Coll, 1993, 14(1): 37-39. (in Chinese with English abstract)
[13]	Takai T, Matsuura S, Nisiho T, et al.Rice yield potenital is closely related to crop growth rate duirng late reproductive peirod.Field Crops Res,2006,96:328-335.
[14]	雷小龙, 刘利, 刘波, 等. 杂交籼稻机械化种植的分蘖特性. 作物学报, 2014, 40(6): 104-1055.
	Lei X L, Liu L, Liu B, et al.Tillering characteristics of indica hybrid rice under mechanized planting.Acta Agron Sin, 2014, 40(6): 104-1055. (in Chinese with English abstract)
[15]	刘利, 雷小龙, 黄光忠, 等. 机械化播栽对杂交稻氮素积累分配及碳氮比的影响. 植物营养与肥料学报, 2014, 20(4): 831-844.
	Liu L, Lei X L, Huang G Z, et al.Influences of mechanical sowing and transplanting on nitrogen accumulation, distribution and C/N of hybrid rice cultivars.J Plant Nutr Fert, 2014, 20(4): 831-844. (in Chinese with English abstract)
[16]	雷小龙, 刘利, 苟文, 等.种植方式对杂交籼稻植株抗倒伏特性的影响. 作物学报, 2013, 39(10): 1814-1825.
	Lei X L, Liu L, Gou w, et al. Effects of planting methods on culm lodging resistance of indica hybrid rjce(Oryza sativa L.).Acta Agron Sin, 2013, 39(10): 1814-1825. (in Chinese with English abstract)
[17]	田青兰, 刘波, 钟晓媛, 等. 不同播栽方式下杂交籼稻非结构性碳水化合物与枝梗和颖花形成及产量性状的关系. 中国农业科学, 2016, 49(1): 35-53.
	Tian Q L, Liu B, Zhong X Y, et al.Relationship of NSC with the formation of branches and spikelets and the yield traits of indica hybrid rice in different planting methods.Sci Agric Sin, 2016, 49(1): 35-53. (in Chinese with English abstract)
[18]	冯永祥, 徐正进, 王聪. 水稻株型的研究进展. 内蒙古民族大学学报:自然科学版, 2003, 18(3): 260-264.
	Feng Y X, Xu Z J, Wang C.Advance on research of rice plant type.J Inner Mon Univ, 2003, 18(3): 260-264. (in Chinese with English abstract)
[19]	艾志勇, 马国辉. 超高产杂交稻茎秆特征与抗倒性关系研究. 杂交水稻, 2010, 25(6): 65-69.
	Ai Z Y, Ma G H.Studies on relationship between culm characteristics and lodging resistance of super-high-yield hybrid rice.Hybrid Rice, 2010, 25(6): 65-69. (in Chinese with English abstract)
[20]	马均, 马文波, 田彦华, 等. 重穗型水稻植株抗倒伏能力的研究. 作物学报, 2004, 30(2): 143-148.
	Ma J, Ma W B, Tian Y H, et al.The culm lodging resistance of heavy panicle type of rice.Acta Agron Sin, 2004, 30(2): 143-148. (in Chinese with English abstract)
[21]	吴晓然, 张巫军, 伍龙梅, 等. 超级杂交籼稻抗倒能力比较及其对氮素的响应. 中国农业科学, 2015, 48(14): 2705-2717.
	Wu X R, Zhang W J, Wu L M, et al, Tang S, Ding C Q, Zhang S H, Ding Y F. Characteristics of lodging resistance of super-hybrid indica rice and its response to nitrogen.Sci Agric Sin, 2015, 48(14): 2705-2717. (in Chinese with English abstract)
[22]	龚振平. 不同营养水平水稻株高及构成变化的研究. 黑龙江农业科学, 1997, 19(6): 18-20.
	Gong Z P.Plant height and its constitutive variation the conditions of different fertilities in rice.Heilongjiang Agric Sci, 1997, 19(6): 18-20.(in Chinese with English abstract)
[23]	凌启鸿, 张洪程, 苏祖芳, 等. 稻作新理论——水稻叶龄模式. 北京: 科学出版社, 1994:19-82.
	Ling Q H, Zhang H C, Su Z F, et al.New Theory of Rice—Rice Leaf Age Model. Beijing: Science Press, 1994, 19-82. (in Chinese with English abstract)
[24]	杨惠杰, 房贤涛, 何花榕, 等. 福建超级稻品种茎秆结构特征及其与抗倒性和产量的关系. 中国生态农业学报, 2012, 20(7): 909-913.
	Yang H J, Fang X T, He H R, et al.Relationship of characteristics of culm construction to lodging resistance and yield of Fujian-bred super-rice cultivars.Chin J Eco-Agric, 2012, 20(7): 909-913. (in Chinese with English abstract)
[25]	赵黎明, 萧长亮, 顾春梅, 等.植物生长调节剂在水稻倒伏上的研究进展. 北方水稻, 2009, 39(3): 114-117.
	Zhao L M, Xiao C L, Gu C M, et al.Research advantage of plant growth regulation on lodging rice.North Rice, 2009, 39(3): 114-117. (in Chinese with English abstract)
[26]	Horie T, Shiraiwa T, Homma K, et al.Can yields of lowland rice resume the increases that they showed in the 1980s?Plant Prod Sci, 2005,8(3): 259-274.
[27]	王亚梁, 张玉屏, 曾研华, 等. 水稻穗形成期高温影响的研究进展. 浙江农业科学, 2014(11): 1681-1685.
	Wang Y L, Zhang Y P, Zeng Y H, et al. Research progress of effect of high temperature on rice panicle formation stage, Zhejiang Agric Sci, 2014(11): 1681-1685. (in Chinese with English abstract)
[28]	姚有礼, 王余龙, 蔡建中. 水稻大穗形成机理的研究(3)品种间每穗颖花现存数与颖花分化和抽穗期物质生产的关系. 江苏农学院学报, 1995, 16(2): 11-16.
	Yao Y L, Wang Y L, Cai J Z.Formation of large panicle in rice (3) varietal difference of survived spikelet number per panicle and its relations with differentiated spikelet number and biomass at heading.Acad J Jiangsu Agric Coll, 1995, 16(2): 11-16. (in Chinese with English abstract)
[29]	Yao Y, Yamamoto Y, Yoshida T, et al.Response of differentiated and degenerated spikelets to top-dressing, shading and day/night temperature treatments in rice cultivars with large panicles.Soil Sci Plant Nutr, 2000, 46(3): 631-641.
[30]	张顺堂, 张桂莲, 陈立云, 等. 高温胁迫对水稻剑叶净光合速率和叶绿素荧光参数的影响. 中国水稻科学, 2011, 25(3): 335-338.
	Zhang S T, Zhang G L, Chen L Y, et al.Effects of high temperature stress on net photosynthetic rate and chlorophyll fluorescence parameters of flag leaf in rice.Chin J Rice Sci, 2011, 25(3): 335-338. (in Chinese with English abstract)
[31]	张桂莲, 陈立云, 张顺堂, 等. 高温胁迫对水稻剑叶保护酶活性和膜透性的影响. 作物学报, 2006, 36(9): 1306-1310.
	Zhang G L, Chen L Y, Zhang S T, et al.Effect of high temperature stress on protective enzyme activities and membrane permeability of flag leaf in rice.Acta Agron Sin, 2006, 36(9): 1306-1310. (in Chinese with English abstract)
[32]	魏金连, 潘晓华. 夜间温度升高对早稻生长发育及产量的影响. 江西农业大学学报, 2008, 30(3): 428-432.
	Pan J L, Pan X H.Effects of night temperature increase on growth and yield of early season rice.Acta Agric Univ Jiangxiensis, 2008, 30(3): 428-432. (in Chinese with English abstract)
[33]	耿立清, 王嘉宇, 陈温福. 孕穗-灌浆期低温对水稻穗部性状的影响. 华北农学报, 2009, 24(3): 107-111.
	Geng L Q, Wang J Y, Chen W F.Effect of low temperature on panicle characters of rice during booting and grain filling period.Acta Agric Boreali-Sin, 2009, 24(3): 107-111. (in Chinese with English abstract)
[34]	杨东, 段留生, 谢华安, 等. 花前光照亏缺对水稻物质积累及生理特性的影响. 中国生态农业学报, 2011, 19(2): 347-352.
	Yang D, Duan L S, Xie H A, Li Z H, Huang T X.Effect of pre-flowering light deficiency on biomass accumulation and physiological characteristics of rice.Chin J Eco-Agric, 2011, 19(2): 347-352. (in Chinese with English abstract)
[35]	谭孟祥, 景元书, 薛杨, 等. 水层深度对早稻有穗分化期遭遇低温过程时叶片生理特性的影响. 中国农业气象, 2015, 36(5): 553-560.
	Tan M X, Jing Y S, Xue Y, et al.Effects of different water depth on leaf physiological characteristics of early rice during panicle primordium suffered to low temperature.Chin J Agrometeorol,2015, 36(5): 553-560. (in Chinese with English abstract)

播栽方式 Planting method	长度 Length/cm			长轴直径 Long axis diameter/mm
播栽方式 Planting method	N1	N2	N3	N1	N2		N3
机直播 MD
宜香优2115 YXY 2115	2.54±0.44 a	7.58±0.79 Aa	10.47±0.42 a	7.02±0.19	7.07±0.12		6.55±0.10
F优498 FY 498	2.03±0.16 b	6.02±0.4 Bb	9.43±0.55 b	6.73±0.01	6.95±0.07		6.47±0.01
机插 MT
宜香优2115 YXY 2115	3.18±0.31 a	8.48±0.14 Aa	9.89±0.65	7.34±0.20	7.36±0.11		6.78±0.08
F优498 FY 498	2.50±0.33 b	6.75±0.41 Bb	10.15±0.35	7.35±0.22	7.36±0.29		6.75±0.22
手插 HT
宜香优2115 YXY 2115	2.96±0.47 a	7.65±0.78 a	10.74±0.24	7.39±0.34 b	7.48±0.17		6.74±0.06
F优498 FY 498	2.45±0.11 a	6.35±0.37 b	10.01±0.71	7.92±0.23 a	7.47±0.14		6.86±0.10
平均值 Mean
机直播 MD	2.29±0.36 Bb	6.80±1.1 Bb	9.95±0.73	6.87±0.2Bc	7.01±0.09 Bb		6.51±0.06 Bb
机插 MT	2.84±0.48 Aa	7.62±1.22 Aa	10.02±0.19	7.34±0.004 Ab	7.36±0.002 ABa		6.76±0.02 Aa
手插 HT	2.71±0.36 Aa	7.00±0.91 Bb	10.38±0.51	7.65±0.37 Aa	7.47±0.004 Aa		6.80±0.08 Aa
宜香优2115 YXY 2115	2.90±0.32 Aa	7.90±0.50 Aa	10.37±0.43 a	7.25±0.20	7.30±0.21		6.69±0.12
F优498 FY 498	2.54±0.44 a	7.58±0.79 Aa	10.47±0.42 a	7.33±0.59	7.26±0.28		6.69±0.20
F值F value
播栽方式 PM	21.05^**	20.41^**	1.54	35.24^**	16.93^*		31.90^**
品种 V	15.87^**	38.48^**	6.32^*	0.86	0.43		0.00
播栽方式×品种 PM×V	0.15	0.26	3.89(^*)	7.33^*	0.35		1.06
播栽方式 Planting method	短轴直径 Short axis diameter/mm			扁平率 Oblate rate/%
播栽方式 Planting method	N1	N2	N3	N1		N2	N3
机直播 MD
宜香优2115 YXY 2115	6.42±0.12 a	6.29±0.04	5.74±0.04	8.53±1.05		11.03±0.93	12.32±0.84 a
F优498 FY 498	6.19±0.06 b	6.24±0.11	5.78±0.10	8.06±0.88		10.10±1.87	10.61±1.35 b
机插 MT
宜香优2115 YXY 2115	6.61±0.19	6.67±0.07	5.98±0.08	9.91±0.95		9.38±1.78	11.69±0.26 a
F优498 FY 498	6.45±0.16	6.63±0.25	6.11±0.25	12.16±0.87		9.88±0.62	9.53±0.69 b
手插 HT
宜香优2115 YXY 2115	6.68±0.19	6.49±0.10	5.89±0.03 b	9.47±1.85 b		13.20±1.34 a	12.54±0.75 Aa
F优498 FY 498	6.75±0.30	6.81±0.26	6.21±0.15 a	14.64±2.76 a		8.72±2.60 b	9.36±0.92 Bb
平均值 Mean
机直播 MD	6.30±0.16 Bb	6.27±0.04 b	5.76±0.03 Bb	8.29±0.34 b		10.56±0.66	11.45±1.21
机插 MT	6.53±0.11 ABa	6.65±0.03 a	6.05±0.09 Aa	11.01±1.59 a		9.63±0.36	10.58±1.53
手插 HT	6.72±0.05 Aa	6.65±0.23 a	6.05±0.23 Aa	11.94±3.65 a		10.86±3.17	10.90±2.25
宜香优2115 YXY 2115	6.57±0.14	6.48±0.19	5.87±0.12	9.30±0.70 b		11.16±1.92	12.18±0.44 Aa
F优498 FY 498	6.46±0.28	6.56±0.29	6.03±0.23	11.47±3.32 a		9.56±0.74	9.83±0.68 Bb
F值F value
播栽方式 PM	15.85^*	13.74^*	16.78^*	10.04^*		0.63	1.52
品种 V	3.06	0.94	5.25(^*)	7.60^*		3.88(^*)	27.86^**
播栽方式×品种 PM×V	2.22	2.27	1.39	3.88(^*)		3.23	1.00
播栽方式 Planting method	外径 Stem diameter/mm			秆型指数 Stem type index
播栽方式 Planting method	N1	N2	N3		N1	N2	N3
机直播 MD
宜香优2115 YXY2115	6.72±0.15 a	6.68±0.08	6.15±0.07		0.270±0.045 b	0.089±0.009 Bb	0.059±0.002 b
F优498 FY498	6.46±0.03 b	6.59±0.07	6.13±0.05		0.320±0.026 a	0.110±0.006 Aa	0.065±0.004 a
机插 MT
宜香优2115 YXY2115	6.98±0.19	7.01±0.06	6.38±0.08		0.220±0.015 b	0.083±0.002 Bb	0.065±0.004
F优498 FY498	6.90±0.18	6.99±0.27	6.43±0.24		0.278±0.032 a	0.104±0.003 Aa	0.063±0.001
手插 HT
宜香优2115 YXY21156.68±0.19	7.04±0.26 b	6.98±0.13	6.32±0.04		0.240±0.027 b	0.092±0.007 Bb	0.059±0.001 b
F优498 FY498	7.34±0.24 a	7.14±0.19	6.53±0.12		0.300±0.024 a	0.113±0.009 Aa	0.066±0.006 a
平均值 Mean
机直播 MD	6.59±0.18 Bc	6.64±0.06 Bb	6.14±0.01 Bb		0.295±0.035 a	0.099±0.015 ab	0.062±0.004
机插 MT	6.94±0.06 ABb	7.00±0.01Aa	6.40±0.04Aa		0.249±0.041 b	0.093±0.015 b	0.064±0.001
手插 HT	7.19±0.21 Aa	7.06±0.11 Aa	6.43±0.15 Aa		0.270±0.042 ab	0.102±0.015 a	0.062±0.005
宜香优2115 YXY2115	6.91±0.17	6.89±0.18	6.28±0.12		0.243±0.025 Bb	0.088±0.005 Bb	0.061±0.003 b
F优498 FY498	6.90±0.44	6.91±0.28	6.36±0.21		0.299±0.021Aa	0.109±0.005Aa	0.065±0.001a
F值F value
播栽方式 PM	30.94^**	19.59^**	24.14^**		5.31(^*)	8.80^*	0.74
品种 V	0.03	0.06	1.75		21.98^**	47.37^**	6.32^*
播栽方式×品种 PM×V	6.04^*	1.13	1.27		0.06	0.00	2.84

播栽方式 Planting method	长度 Length/cm			长轴直径 Long axis diameter/mm
播栽方式 Planting method	N1	N2	N3	N1	N2		N3
机直播 MD
宜香优2115 YXY 2115	2.54±0.44 a	7.58±0.79 Aa	10.47±0.42 a	7.02±0.19	7.07±0.12		6.55±0.10
F优498 FY 498	2.03±0.16 b	6.02±0.4 Bb	9.43±0.55 b	6.73±0.01	6.95±0.07		6.47±0.01
机插 MT
宜香优2115 YXY 2115	3.18±0.31 a	8.48±0.14 Aa	9.89±0.65	7.34±0.20	7.36±0.11		6.78±0.08
F优498 FY 498	2.50±0.33 b	6.75±0.41 Bb	10.15±0.35	7.35±0.22	7.36±0.29		6.75±0.22
手插 HT
宜香优2115 YXY 2115	2.96±0.47 a	7.65±0.78 a	10.74±0.24	7.39±0.34 b	7.48±0.17		6.74±0.06
F优498 FY 498	2.45±0.11 a	6.35±0.37 b	10.01±0.71	7.92±0.23 a	7.47±0.14		6.86±0.10
平均值 Mean
机直播 MD	2.29±0.36 Bb	6.80±1.1 Bb	9.95±0.73	6.87±0.2Bc	7.01±0.09 Bb		6.51±0.06 Bb
机插 MT	2.84±0.48 Aa	7.62±1.22 Aa	10.02±0.19	7.34±0.004 Ab	7.36±0.002 ABa		6.76±0.02 Aa
手插 HT	2.71±0.36 Aa	7.00±0.91 Bb	10.38±0.51	7.65±0.37 Aa	7.47±0.004 Aa		6.80±0.08 Aa
宜香优2115 YXY 2115	2.90±0.32 Aa	7.90±0.50 Aa	10.37±0.43 a	7.25±0.20	7.30±0.21		6.69±0.12
F优498 FY 498	2.54±0.44 a	7.58±0.79 Aa	10.47±0.42 a	7.33±0.59	7.26±0.28		6.69±0.20
F值F value
播栽方式 PM	21.05^**	20.41^**	1.54	35.24^**	16.93^*		31.90^**
品种 V	15.87^**	38.48^**	6.32^*	0.86	0.43		0.00
播栽方式×品种 PM×V	0.15	0.26	3.89(^*)	7.33^*	0.35		1.06
播栽方式 Planting method	短轴直径 Short axis diameter/mm			扁平率 Oblate rate/%
播栽方式 Planting method	N1	N2	N3	N1		N2	N3
机直播 MD
宜香优2115 YXY 2115	6.42±0.12 a	6.29±0.04	5.74±0.04	8.53±1.05		11.03±0.93	12.32±0.84 a
F优498 FY 498	6.19±0.06 b	6.24±0.11	5.78±0.10	8.06±0.88		10.10±1.87	10.61±1.35 b
机插 MT
宜香优2115 YXY 2115	6.61±0.19	6.67±0.07	5.98±0.08	9.91±0.95		9.38±1.78	11.69±0.26 a
F优498 FY 498	6.45±0.16	6.63±0.25	6.11±0.25	12.16±0.87		9.88±0.62	9.53±0.69 b
手插 HT
宜香优2115 YXY 2115	6.68±0.19	6.49±0.10	5.89±0.03 b	9.47±1.85 b		13.20±1.34 a	12.54±0.75 Aa
F优498 FY 498	6.75±0.30	6.81±0.26	6.21±0.15 a	14.64±2.76 a		8.72±2.60 b	9.36±0.92 Bb
平均值 Mean
机直播 MD	6.30±0.16 Bb	6.27±0.04 b	5.76±0.03 Bb	8.29±0.34 b		10.56±0.66	11.45±1.21
机插 MT	6.53±0.11 ABa	6.65±0.03 a	6.05±0.09 Aa	11.01±1.59 a		9.63±0.36	10.58±1.53
手插 HT	6.72±0.05 Aa	6.65±0.23 a	6.05±0.23 Aa	11.94±3.65 a		10.86±3.17	10.90±2.25
宜香优2115 YXY 2115	6.57±0.14	6.48±0.19	5.87±0.12	9.30±0.70 b		11.16±1.92	12.18±0.44 Aa
F优498 FY 498	6.46±0.28	6.56±0.29	6.03±0.23	11.47±3.32 a		9.56±0.74	9.83±0.68 Bb
F值F value
播栽方式 PM	15.85^*	13.74^*	16.78^*	10.04^*		0.63	1.52
品种 V	3.06	0.94	5.25(^*)	7.60^*		3.88(^*)	27.86^**
播栽方式×品种 PM×V	2.22	2.27	1.39	3.88(^*)		3.23	1.00
播栽方式 Planting method	外径 Stem diameter/mm			秆型指数 Stem type index
播栽方式 Planting method	N1	N2	N3		N1	N2	N3
机直播 MD
宜香优2115 YXY2115	6.72±0.15 a	6.68±0.08	6.15±0.07		0.270±0.045 b	0.089±0.009 Bb	0.059±0.002 b
F优498 FY498	6.46±0.03 b	6.59±0.07	6.13±0.05		0.320±0.026 a	0.110±0.006 Aa	0.065±0.004 a
机插 MT
宜香优2115 YXY2115	6.98±0.19	7.01±0.06	6.38±0.08		0.220±0.015 b	0.083±0.002 Bb	0.065±0.004
F优498 FY498	6.90±0.18	6.99±0.27	6.43±0.24		0.278±0.032 a	0.104±0.003 Aa	0.063±0.001
手插 HT
宜香优2115 YXY21156.68±0.19	7.04±0.26 b	6.98±0.13	6.32±0.04		0.240±0.027 b	0.092±0.007 Bb	0.059±0.001 b
F优498 FY498	7.34±0.24 a	7.14±0.19	6.53±0.12		0.300±0.024 a	0.113±0.009 Aa	0.066±0.006 a
平均值 Mean
机直播 MD	6.59±0.18 Bc	6.64±0.06 Bb	6.14±0.01 Bb		0.295±0.035 a	0.099±0.015 ab	0.062±0.004
机插 MT	6.94±0.06 ABb	7.00±0.01Aa	6.40±0.04Aa		0.249±0.041 b	0.093±0.015 b	0.064±0.001
手插 HT	7.19±0.21 Aa	7.06±0.11 Aa	6.43±0.15 Aa		0.270±0.042 ab	0.102±0.015 a	0.062±0.005
宜香优2115 YXY2115	6.91±0.17	6.89±0.18	6.28±0.12		0.243±0.025 Bb	0.088±0.005 Bb	0.061±0.003 b
F优498 FY498	6.90±0.44	6.91±0.28	6.36±0.21		0.299±0.021Aa	0.109±0.005Aa	0.065±0.001a
F值F value
播栽方式 PM	30.94^**	19.59^**	24.14^**		5.31(^*)	8.80^*	0.74
品种 V	0.03	0.06	1.75		21.98^**	47.37^**	6.32^*
播栽方式×品种 PM×V	6.04^*	1.13	1.27		0.06	0.00	2.84

播栽方式与品种 PM and V	高效叶面积率 HELR/%	叶面积指数 LAA	颖花叶面积比 RSLA/(粒·cm^-2)	实粒叶面积比 RFSLA/(粒·cm^-2)
机直播 MD
宜香优2115 YXY2115	68.85±1.34	9.92±0.19 a	0.34±0.03 Bb	0.31±0.01 Bb
F优498 FY498	68.99±1.71	8.58±1.63 b	0.55±0.06 Aa	0.51±0.06 Aa
机插 MT
宜香优2115 YXY2115	64.36±0.94	9.43±0.62	0.41±0.04 Bb	0.35±0.05 Bb
F优498 FY498	62.37±6.11	9.10±0.41	0.58±0.03 Aa	0.53±0.03 Aa
手插 HT
宜香优2115 YXY2115	66.80±1.93	9.70±0.24 Aa	0.38±0.04 Bb	0.33±0.04 Bb
F优498 FY498	67.53±5.77	6.93±0.21 Bb	0.68±0.05 Aa	0.63±0.04 Aa
平均值 Mean
机直播 MD	68.92±0.10	9.25±0.94	0.44±0.15 b	0.41±0.15
机插 MT	63.37±1.41	9.26±0.23	0.49±0.12 ab	0.44±0.13
手插 HT	67.17±0.52	8.32±1.96	0.53±0.22 a	0.48±0.21
宜香优2115 YXY2115	66.68±2.25	9.68±0.25 Aa	0.37±0.03 Bb	0.33±0.02 Bb
F优498 FY498	66.33±3.48	8.21±1.13 Bb	0.6±0.07 Aa	0.56±0.06 Aa
F值F value
播栽方式PM	3.79	2.75	5.39(^*)	3.42
品种 V	0.06	19.36^**	150.05^**	209.07^**
播栽方式×品种PM×V	0.29	4.47(^*)	4.31(^*)	5.62^*
播栽方式与品种 PM and V	粒重叶面积比 RGWLA /(mg·cm^-2)	单穗实粒重 GWPP /g	穗长 LP /cm	着粒密度 GD /(粒·cm^-1)
机直播 MD
宜香优2115 YXY2115	10.28±0.33 Bb	3.89±0.14 Bb	25.47±0.48 a	5.03±0.28 Bb
F优498 FY498	14.96±1.76 Aa	5.26±0.54 Aa	25.11±0.71 a	7.71±0.94 Aa
机插MT
宜香优2115 YXY2115	11.95±1.44 b	4.51±0.44 Bb	28.74±0.14 Aa	5.33±0.32 Bb
F优498 FY498	15.06±1.13a	5.78±0.72 Aa	26.84±0.37 Bb	8.26±0.47 Aa
手插 HT
宜香优2115 YXY2115	11.15±1.31 Bb	4.27±0.27 Bb	27.48±0.32 a	5.26±0.18 Bb
F优498 FY498	18.6±1.41 Aa	6.03±0.11 Aa	26.29±0.61 b	8.41±0.16 Aa
平均值 Mean
机直播 MD	12.62±3.31 b	4.58±0.97	25.29±0.25 Bc	6.37±1.89
机插 MT	13.51±2.2 ab	5.15±0.90	27.79±1.35 Aa	6.79±2.08
手插 HT	14.87±5.27 a	5.15±1.25	26.88±0.84 Ab	6.84±2.23
宜香优2115 YXY2115	11.13±0.83 Bb	4.23±0.31 Bb	27.23±1.65 Aa	5.21±0.16 Bb
F优498 FY498	16.21±2.07 Aa	5.69±0.39 Aa	26.08±0.88 Bb	8.13±0.37 Aa
F值F value
播栽方式PM	4.13	5.39(^*)	47.19^**	2.09
品种 V	86.39^**	150.05^**	22.17^**	171.41^**
播栽方式×品种PM×V	5.38^*	4.31(^*)	3.36	0.38

播栽方式与品种 PM and V	高效叶面积率 HELR/%	叶面积指数 LAA	颖花叶面积比 RSLA/(粒·cm^-2)	实粒叶面积比 RFSLA/(粒·cm^-2)
机直播 MD
宜香优2115 YXY2115	68.85±1.34	9.92±0.19 a	0.34±0.03 Bb	0.31±0.01 Bb
F优498 FY498	68.99±1.71	8.58±1.63 b	0.55±0.06 Aa	0.51±0.06 Aa
机插 MT
宜香优2115 YXY2115	64.36±0.94	9.43±0.62	0.41±0.04 Bb	0.35±0.05 Bb
F优498 FY498	62.37±6.11	9.10±0.41	0.58±0.03 Aa	0.53±0.03 Aa
手插 HT
宜香优2115 YXY2115	66.80±1.93	9.70±0.24 Aa	0.38±0.04 Bb	0.33±0.04 Bb
F优498 FY498	67.53±5.77	6.93±0.21 Bb	0.68±0.05 Aa	0.63±0.04 Aa
平均值 Mean
机直播 MD	68.92±0.10	9.25±0.94	0.44±0.15 b	0.41±0.15
机插 MT	63.37±1.41	9.26±0.23	0.49±0.12 ab	0.44±0.13
手插 HT	67.17±0.52	8.32±1.96	0.53±0.22 a	0.48±0.21
宜香优2115 YXY2115	66.68±2.25	9.68±0.25 Aa	0.37±0.03 Bb	0.33±0.02 Bb
F优498 FY498	66.33±3.48	8.21±1.13 Bb	0.6±0.07 Aa	0.56±0.06 Aa
F值F value
播栽方式PM	3.79	2.75	5.39(^*)	3.42
品种 V	0.06	19.36^**	150.05^**	209.07^**
播栽方式×品种PM×V	0.29	4.47(^*)	4.31(^*)	5.62^*
播栽方式与品种 PM and V	粒重叶面积比 RGWLA /(mg·cm^-2)	单穗实粒重 GWPP /g	穗长 LP /cm	着粒密度 GD /(粒·cm^-1)
机直播 MD
宜香优2115 YXY2115	10.28±0.33 Bb	3.89±0.14 Bb	25.47±0.48 a	5.03±0.28 Bb
F优498 FY498	14.96±1.76 Aa	5.26±0.54 Aa	25.11±0.71 a	7.71±0.94 Aa
机插MT
宜香优2115 YXY2115	11.95±1.44 b	4.51±0.44 Bb	28.74±0.14 Aa	5.33±0.32 Bb
F优498 FY498	15.06±1.13a	5.78±0.72 Aa	26.84±0.37 Bb	8.26±0.47 Aa
手插 HT
宜香优2115 YXY2115	11.15±1.31 Bb	4.27±0.27 Bb	27.48±0.32 a	5.26±0.18 Bb
F优498 FY498	18.6±1.41 Aa	6.03±0.11 Aa	26.29±0.61 b	8.41±0.16 Aa
平均值 Mean
机直播 MD	12.62±3.31 b	4.58±0.97	25.29±0.25 Bc	6.37±1.89
机插 MT	13.51±2.2 ab	5.15±0.90	27.79±1.35 Aa	6.79±2.08
手插 HT	14.87±5.27 a	5.15±1.25	26.88±0.84 Ab	6.84±2.23
宜香优2115 YXY2115	11.13±0.83 Bb	4.23±0.31 Bb	27.23±1.65 Aa	5.21±0.16 Bb
F优498 FY498	16.21±2.07 Aa	5.69±0.39 Aa	26.08±0.88 Bb	8.13±0.37 Aa
F值F value
播栽方式PM	4.13	5.39(^*)	47.19^**	2.09
品种 V	86.39^**	150.05^**	22.17^**	171.41^**
播栽方式×品种PM×V	5.38^*	4.31(^*)	3.36	0.38

器官 Organ	机直播 MD	机插 MT	手插 HT	宜香优2115 YXY2115	F优498 F you 498	F值 F value
器官 Organ	机直播 MD	机插 MT	手插 HT	宜香优2115 YXY2115	F优498 F you 498	播栽方式 PM	品种 V	播栽方式×品种 PM×V
叶 Leaf
24DBH	1.06±0.14	0.93±0.04	1.11±0.38	0.92±0.07Bb	1.15±0.24Aa	3.57	39.03^**	22.48^**
20DBH	1.15±0.17 Aa	1.06±0.01 Bb	1.12±0.08 ABa	1.05±0.02 Bb	1.17±0.11 Aa	11.74^*	15.17^**	5.95^*
16DBH	1.07±0.12	1.17±0.09	1.22±0.04	1.17±0.07	1.14±0.13	2.06	4.67(^*)	33.48^**
12DBH	1.11±0.06 b	1.37±0.14 a	1.28±0.15 ab	1.20±0.07	1.31±0.21	8.40^*	2.88	2.14
8DBH	1.11±0.04 Bb	1.44±0.05 Aa	1.38±0.09 ABa	1.31±0.17	1.31±0.20	14.3^*	0.001	2.06
4DBH	1.28±0.05 b	1.39±0.23 ab	1.55±0.29 a	1.54±0.22 Aa	1.27±0.07 Bb	4.59(^*)	30.43^**	4.37(^*)
0DBH	1.38±0.03	1.62±0.07	1.59±0.13	1.59±0.15	1.48±0.11	2.93	2.26	0.32
叶鞘 Sheath
24DBH	0.95±0.02	0.92±0.02	0.84±0.11	0.88±0.10	0.93±0.01	1.35	2.21	2.51
20DBH	1.05±0.03	1.06±0.01	1.05±0.09	1.02±0.03b	1.08±0.03a	0.35	8.05^*	2.25
16DBH	1.11±0.01	1.16±0.06	1.19±0.07	1.12±0.02	1.18±0.07	0.28	2.99	1.04
12DBH	1.12±0.1 b	1.43±0.14 a	1.26±0.21 ab	1.17±0.15 Bb	1.38±0.17 Aa	10.44^*	35.91^**	1.65
8DBH	1.15±0.04 Cc	1.56±0.03 Aa	1.43±0.16 Bb	1.34±0.23	1.42±0.21	167.07^**	3.01	2.66
4DBH	1.44±0.04	1.60±0.16	1.62±0.19	1.63±0.19 a	1.48±0.01 b	2.67	7.90^*	3.60(^*)
0DBH	1.65±0.14 b	1.88±0.09 a	1.76±0.04 ab	1.83±0.10	1.70±0.14	7.81 ^*	2.81	0.35
茎秆 Stem
24DBH	0.31±0.06 Bb	0.39±0.02 Aa	0.36±0.03 ABa	0.37±0.03	0.34±0.06	16.67^*	3.02	4.92(^*)
20DBH	0.49±0.02	0.53±0.10	0.49±0.08	0.50±0.09	0.50±0.04	1.01	0.02	9.24^*
16DBH	0.64±0.04 Bb	0.78±0.18 Aa	0.59±0.03 Bb	0.71±0.18 Aa	0.63±0.05 Bb	26.58^**	63.81^**	86.14^**
12DBH	0.82±0.12 Bb	1.05±0.07 Aa	0.85±0.11 Bb	0.98±0.11 Aa	0.84±0.14 Bb	17.28^*	32.77^**	0.79
8DBH	0.99±0.08 Bc	1.24±0.20 Aa	1.11±0.003 Bb	1.18±0.18 a	1.05±0.09 b	45.97^**	12.39^*	4.73(^*)
4DBH	1.23±0.05 Bb	1.39±0.24 Aa	1.28±0.13 Bb	1.40±0.15 Aa	1.20±0.02 Bb	32.03^**	15.51^**	2.25
0DBH	1.42±0.25 Bb	1.68±0.21 Aa	1.59±0.09 Aa	1.69±0.12 a	1.44±0.16 b	28.27^**	12.43^*	0.89